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Ox at Mn1, 2.1 Å from O5, suggests that Ox either forms the O–O bond with O5 in the S3→S0 transition (case 1 in Fig. 5), or that Ox is placed between Ca and Mn1 to replace O5 during O2 formation or release. In the latter case, O–O bond formation may occur between O5 and W2 (2) or O5 and W3 (3). 

Read more: diamond.ac.uk/Science/Research/Highlights/2019/timely-solution-for-photosynthetic-oxygen-evolving-clock.html

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